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Co-phylogenetic Analysis Tool

CopyCat provides an easy and fast access to cophylogenetic analyses. It incorporates a wrapper for the program ParaFit, which conducts a statistical test for the presence of congruence between host and parasite phylogenies.

CopyCat offers various features, such as the creation of customized host-parasite association data and the computation of phylogenetic host/parasite trees based on the NCBI taxonomy

Use of the program is free for academic purposes at an academic institute. For all other uses, please contact the authors.


If you use CopyCat, please cite the following paper: Jan P. Meier-Kolthoff, Alexander F. Auch, Daniel H. Huson, Markus Göker. COPYCAT: Co-phylogenetic Analysis tool. Bioinformatics, 23(7):898-900, 2007. PDF

Supplemental material:

Users of AxParafit and AxPcoords may also cite: Alexandros Stamatakis, Alexander F. Auch, Jan P. Meier-Kolthoff, Markus Göker. AxPcoords & parallel AxParafit: statistical co-phylogenetic analyses on thousands of taxa. BMC Bioinformatics 8:405, 2007. PDF


copycat_new  New version 2.0 available on new Copycat website:

Version 1.14, including AxParafit and AxPcoords which are highly optimized versions of Pierre Legendre's Parafit and DistPCoA programs:

Older versions can be found here.

These installers do not contain a Java RunTime Environment (JRE). Please make sure that you have one installed, version 1.5.0 or higher.


To find out more about the program, take a look at the user manual.


The program was written by Jan P. Meier-Kolthoff, Alexander F. Auch, Daniel H. Huson, and Markus Göker.

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